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雌蕊先熟的

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Inflorescence spikelike, cylindrical, composed of spiny or bristly deciduous burrs arranged along an angular, often sinuous rachis; burrs sessile or with an obconical basal stipe, each composed of 1 or more sessile spikelets surrounded by an involucre of spines and bristles; bristles flexuous or more often spinous,± flattened, grooved on the outer face, united below, the degree of union varying from a small basal disk to a deep cupule, inner spines or bristles often ciliate around spikelets. Spikelets lanceolate, acute; glumes unequal, shorter than spikelet, lower sometimes suppressed; lower floret membranous, staminate or neuter; upper floret firmer, protogynous.

花序穗状,圆筒状,由落叶的毛刺沿着一具角安排的具刺的或具刚毛,通常波状的轴组成;毛刺无梗或具基部的倒锥状柄,每一个由1无梗或更多组成以总苞的刺和刚毛包围;或更多的刚毛曲折通常spinous,多少扁平,在外面上具凹槽,下面的合生,协会的度从一小的基部的花盘变化到一深的壳斗,小穗披针形,锐尖;不等长的颖片,短于小穗,降低有时压抑;更低的小花膜质,雄蕊花的或中性;坚定的上面小花,雌蕊先熟。

Spikelets protogynous, strongly laterally compressed, floret 1, falling entire from pedicel; rachilla extension absent; glumes equal,± equaling and enclosing floret, broadly lanceolate to oblong, membranous to thinly leathery, prominently 3-veined, strongly keeled, keel usually ciliate, infrequently winged, lower margins almost free or connate for up to half their length, apex obtuse, acute or shortly awned; lemma broadly lanceolate to ovate, usually thinly membranous, keeled, obscurely 5-veined, smooth, glabrous, lower margins often connate, awned from lower back, apex truncate to acute; awn straight when short or geniculate when longer, column smooth, usually twisted at maturity, bristle scabrid; palea absent or very small.

雌蕊先熟的小穗,强烈侧面压扁,小花1,从花梗那里变为全缘;小穗轴扩展缺席;颖片等长,多少等长和包围小花,宽披针形的到长圆形,膜质到薄革质,显著3脉,强烈龙骨状,外稃宽披针形的到卵形,直的芒什么时候短或者膝曲长时间,顺利的柱子,通常成熟时扭曲,粗糙的硬毛;内稃缺席或者很小。

Protogyny The maturation of the stigma before the anthers.

雌蕊先熟:指柱头先于花粉囊成熟的现象。

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