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胼胝质

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Pollen morphology of the Chinese monotypic genus Dichotomanthes and its out-group Prinsepia utilis were investigated from the point of view of pollen development. The SEM observation revealed that there is not any change in morphology and sculpture of pollen grains from the young pollen grain seperated from callose of tetraspore to the mature pollen grain in Dichotomanthes tristaniaecarpa. Only the volume of pollen grain increased with maturity.

从发育的角度研究了中国特有单种属Dichotomanthes Kurz及与其系统学研究有关的外类群Prinsepia utilis的花粉形态,扫描电镜观察显示Dichotomanthes花粉粒自脱离四分体胼胝质膜开始至成熟二核花粉粒不同发育时期,花粉形态和外壁纹饰未见变化,仅花粉体积随成熟度增加而有所增大。

Results there were unconsciousness, respiratory rhythm disturbance and hyporeflexia of pupil light reflex immediately after injury, and reactiveness decrease and activity retardation still existed even after resuming consciousness. at anatomical scene, subarachnoid hemorrhage or cerebroventricular haemorrhage were widespread. at an early stage, there were swelling, collapse, and axonal retraction ball formation at cortico-medulla junction, callosum, brainstem, and cerebellar white matter under microscope. but at the later stage, gitter cell proliferation and nest-like aggregation were major pathophysiological changes at focal brain tissue.

结果 伤后大鼠均即刻出现原发昏迷,其中2只于损伤后20 min内死亡,余持续时间1-30 min不等;伤后大鼠呼吸节律紊乱,瞳孔对光反射减弱或消失,醒后均有程度不等的反应性下降,肢体活动迟缓;肉眼可见广泛蛛网膜下腔出血或脑室出血;光镜下可见皮髓交界区、胼胝体区、脑干、小脑白质等部位的神经轴索有不同程度的肿胀、断裂、轴索球形成,后期有小胶质细胞增生,局部呈巢样聚集。

Spikelets similar, lanceolate, dorsally compressed; callus bearded with hairs shorter than, as long as, or longer than the spikelet; glumes papery or membranous; lower floret usually represented by a hyaline sterile lemma; upper floret bisexual, lemma hyaline, awned or awnless.

小穗相似,披针形,背压扁;胼胝体具髯毛的具毛短于,倍于,比小穗或更长;颖片纸质的或膜质;通常被一透明的不育的外稃描述的更低的小花;上面小花阴阳人,外稃透明,具芒的或awnless。

Spikelets 3–4 mm, brown; callus hairs a little shorter than spikelet, brown; lower glume oblong-lanceolate, papery becoming herbaceous toward apex, back brown-hirsute, apex narrowly truncate; lower lemma 3/4 length of glumes; upper lemma oblong, obtuse and awnless or minutely mucronate, varying to shortly 2-toothed and awned from sinus; awn twisted, bent, up to 6 mm.

小穗3-4毫米,棕色;胼胝体毛一小短于小穗,棕色;狭的更低的颖片长圆状披针形,纸质的变得的草质的向先端,背面棕色具粗毛,先端截形;更低的外稃3/4段颖片;上面外稃长圆形,短尖的钝和awnless或微小的,变化随着短2齿和从那里凹缺;芒扭曲,弯曲,可达6毫米花粉囊3,1.6-2.2毫米。

Racemes (4–)10–40(–100),(8–)10–30 cm; rachis internodes glabrous, scaberulous or smooth, nodes glabrous; lower pedicel 0.5–1.5 mm, upper pedicel 1.5–4 mm. Spikelets 4–6.5 mm, pilose or glabrous, awned; callus hairs 5–8 mm, exceeding the spikelet; glumes subequal, membranous, 4–6.5 mm, 5-veined, back glabrous or pilose, puberulous at apex and along upper margins, apex acuminate; lower lemma lanceolate, hyaline, 3.5–4 mm, veinless, apex and margins puberulous, otherwise glabrous; upper lemma similar to lower, 2.5–3.5 mm; awn geniculate, 4–12 mm; upper palea a 1–2 mm scale.

总状花序(4-)10-40(-100),(8-)10-30厘米;轴节间的无毛,微糙的或平滑,节点无毛;更低的花梗0.5-1.5毫米,上面花梗1.5-4毫米小穗4-6.5毫米,具柔毛或无毛,具芒;胼胝体毛5-8毫米,超过小穗;颖片近等长,膜质,毫米,5-脉,背无毛或具柔毛,被微柔毛在先端和沿着上面边缘,先端渐尖;披针形,透明的下部外稃,3.5-4毫米,无脉,被微柔毛的先端和边缘,或无毛;类似于上面外稃降低,2.5-3.5毫米;膝曲的芒,4-12毫米;上面内稃一1-2毫米鳞片。

Sessile spikelet subterete; callus long, pungent, bearded; lower glume leathery, obtuse; upper glume awnless; lower floret reduced to a hyaline lemma; upper lemma stipitiform, entire, passing into a stout geniculate awn with hairy column.

无梗小穗近圆柱状;胼胝体长,锐尖,具髯毛;更低的颖片革质,钝;上面颖片awnless;更低的小花退化至一透明的外稃;膝曲的柄状,全缘,度过的上面外稃成为一粗壮具有毛柱状。

Spikelets laterally compressed, florets 2, lower floret fertile, upper floret reduced to a small empty lemma, disarticulating above glumes; glumes about as long as florets, broad, membranous, 1-veined; lower glume awnless; upper glume with an oblique, stout, subapical awn; callus pubescent; lemma of fertile floret dark brown, keeled, broad, cartilaginous to leathery, 3-veined, hairy on veins, apex acute to emarginate, awnless or with a fine, subapical awn-point; palea nearly as long as lemma, keels scabrous or ciliolate; upper floret cuneate or obovate, glabrous.

小穗侧面压扁,小花2,能育,上面的更下部的小花退化至小空外稃,高于颖片脱节;关于倍于小花的颖片,宽,膜质,1脉;更低的颖片awnless;上面颖片具一偏斜,粗壮,芒;胼胝体短柔毛;暗褐色的肥沃的小花的外稃,翻转,宽,软骨的到革质,3脉,内稃近等长外稃,龙骨粗糙或者具短缘毛;上面小花楔形的或倒卵形,无毛。

Spikelets lanceolate to narrowly oblong, dorsally compressed; callus short, bearded, hairs usually less than 1/2 spikelet length; glumes cartilaginous to leathery; lower glume flat or slightly concave, hairy, back veinless or inconspicuously veined, flanks rounded in lower half, keeled toward apex, acute to truncate; upper glume boat-shaped, 1-keeled down midline; lower floret reduced to an empty hyaline lemma, rarely absent; upper lemma linear to oblong, occasionally broader, shortly 2-toothed or 2-lobed to middle, awned E.

小穗披针形的到狭长圆形,背压扁;胼胝体短,具髯毛,不到1/2的小穗长度的通常毛;颖片软骨质的到革质;更下部的颖片平或稍凹,有毛,背无脉或不明显,侧面圆形在下半部,龙骨状向先端,锐尖到截形;舟状的上面颖片,沿着midline龙骨状的1;更低的小花退化至一空的透明的外稃,很少无;上面外稃线形到长圆形,偶有宽,2齿或2裂的至中部,具芒的E。

Sessile spikelet 3.5–4.5 mm, pallid; callus hairs 1/8 spikelet length; lower glume oblong-lanceolate, membranous, back slightly concave, villous except near apex, veinless between keels, upper keels smooth or pectinate-scabrid, apex narrowly truncate or 2-toothed; upper glume apex with 2–2.5 mm awn; upper lemma narrow, 2-toothed in upper 1/4; awn ca.

无梗小穗3.5-4.5毫米,灰白色;胼胝体毛1/8的小穗长度;长圆状披针形的下部的颖片,膜质,背稍凹,除近先端之外的具长柔毛,上面颖片先端具2-2.5毫米芒;在上面1/4里狭窄,2齿状的上面外稃;芒约1厘米,柱子具短缘毛。

There is no free microspores released fromthe tetrads and no pollen grains were formed in mutant anther.

这一结果表明MS157 基因可能通过胼胝质降解途径,对雄配子的发育进行调控。

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