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Leaves sessile; blade broadly oblong or oblong-ovate to narrowly oblong or oblong-lanceolate, 2-6.5(-8)× 1-3.2 cm, thickly papery, abaxially rather paler but not glaucous; laminar glands dense, unequal dots or short streaks; intramarginal glands dense; main lateral veins 4-6-paired, with densely reticulate tertiary venation not or scarcely prominent abaxially; base cordate-amplexicaul, apex rounded to shallowly retuse.

叶无柄 叶片宽长圆形或长圆状卵形到狭长圆形或长圆状披针形, 2-6.5(-8)* 1-3.2 厘米,厚纸质,背面相当浅但不是苍白;片状腺体浓,不相等的小圆点或者短的条纹;近边缘内腺体浓; 4-6配对的主要侧脉,突出的具密网状第三脉不或几乎不;抱茎基部心形,先端圆形对浅微凹。

Dioecious monoecious, reproductive structures axillary, solitary. Pollen cones pedicellate, globose, with overlapping bracts at base; microsporophylls 6-14, shield-shaped, each with 4-9 radially arranged pollen sacs. Seed-bearing structures subsessile, with several overlapping bracts basically. Ovule erected, entrusted with disc-like embryophore. After fertilization, embryophore developed into fleshy, cup-like and red aril.

雌雄异株,球花单生于叶腋,有短柄;雄球花圆球形,基部覆瓦状排列的苞片,雄蕊6-14枚,盾状,花药4-9丝,辐射状排列;雌球花几乎无柄,基部有多数覆瓦状排列的苞片,胚珠直立,基部托以圆盘状的珠托,受精后珠托发育成肉质、杯状、红色的假种皮。

Fruit dehiscent siliques, linear, latiseptate, sessile; valves with a prominent midvein, glabrous or rarely pubescent with stalked, forked and 3-rayed trichomes, smooth; replum rounded; septum complete; style obsolete or to 0.2 mm; stigma capitate, entire.

果开裂长角果,线形,,无梗;裂爿具突出中脉,无毛或很少短柔毛具柄,分叉和伞辐毛状体,平滑;假隔膜绕行;隔膜完全;花柱废退的或到0.2毫米;柱头头状,全缘。

Stamens numerous; anthers elliptic to oblong; filaments linear. Follicles 5--40, sessile, sometimes stipitate, with branching transverse veins, styles distinct or nearly absent; ovules several to many.

花瓣无;雄蕊多;花药椭圆形到长圆形;花丝线形蓇葖果5-40,无梗,有时具柄,具分枝横向脉,花柱离生的或近无;胚珠数枚到多数。

Male flowers 4-merous, sessile; perianth lobes elliptic, ca. 1 mm, strigillose, connate at base. Fruiting perianth rhomboid-obovoid, compressed, ca. 1.2 mm, smooth, strigillose on shoulder, base cuneate or stipitate, apex with short neck, 2-toothed.

雄花4瓣,无梗;花被裂片椭圆形,约1毫米,疏生微具短糙伏毛,在基部合生;雌花花被斜方形倒卵球形,压扁的,约1.2毫米的,平滑,具短糙伏毛在肩,基部楔形或具柄的,先端具短颈,2齿上。

Leaves subsessile but not clasping; petals 5-8 mm; stigma surrounded by anthers of longer stamens at anthesis.

叶近无柄但不抱茎;花瓣5-8毫米;长的雄蕊在花期的花药包围的柱头。

Basal petioles 1.5–5 cm, sheaths long, broad; blade oblong, 2–4 × 1–2 cm, pinnate; pinnae sessile,(3–) 4–6 pairs, remote; ultimate segments long-ovate to suborbicular, 5–10 × 3–8 mm, base subtruncate or rounded, entire or apex shallowly 1–3-toothed, sometimes 3–4-crenate; terminal pinnae obovate or orbicular, base cuneate, apex more deeply 3-lobed.

基部的叶柄1.5-5厘米,鞘长,宽;叶片长圆形, 2-4 * 1-2 厘米,羽状;羽片无柄,(3-)4-6 对,遥远;末回裂片长卵形到近圆形, 5-10 * 3-8 毫米,基部近截形或圆形,有时,浅全缘或先端1-3齿状具圆齿;更深的顶生的羽片倒卵形的或圆形,基部楔形,先端3浅裂。

Cauline leaves petiolate or sessile and cuneate or auriculate, pinnately or palmately veined, entire, dentate, or crenate; ultimate veins ending or not with apiculate callosities.

具叶柄或者无柄和楔形的茎生叶或者耳形,成羽状或者掌状脉,整个,有齿,或具圆齿;末级的脉以具细尖的胼胝体结束不的或。

Corolla rotate; filaments connate basally into a ring or thin tube; leaves sessile or only lower ones short petiolate (except in white-flowered species).

花冠辐状;基部的花丝合生成为一环或者薄的筒部;叶无柄或只最下部的短叶柄。

Stems 18-100(-160) cm tall, well-branched above, densely gray villous on lower part, mixed above with short glandular hairs, often with raised lines decurrent from margins of petioles. Leaves subsessile or lower ones with petioles 1-3 mm; cauline blade lanceolate-elliptic to narrowly lanceolate or oblong-lanceolate, 3-12 × 0.5-2.5 cm, both surfaces villous, base usually rounded, margin denticulate with 15-60 teeth per side, apex subacute.

茎18-100(-160)厘米高,在上面的多分枝,叶近无柄或下部的具叶柄1-3毫米;茎生叶披针形椭圆的到狭披针形或者长圆状披针形, 3-12 * 0.5-2.5 两面,厘米具长柔毛,基部通常圆形,边缘具小齿具齿每边,近尖的先端。

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