托叶的

The distribution of the qualitative characters The performance of the 39 chief qualitative characters were observed and recorded,according to the test method of morphological characters in the .The results of distribution proportion of each character in cultivars group showed that the distribution of 32 characters(number of ray flower rings,floret curliness,shape of leaf base,size of stipule and so on) were very asymmetrical with near or above 70%cultivars concentrated upon one or two performance, however,the distribution of the other 7 characters(floret type,surface color of outer floret,back color of outer floret,color of disc floret,claw in base of sinus between lateral lobes,stem erectness and stem color) in the cultivars group were relatively symmetrical.

质量性状分布参照《菊花DUS测试指南》的形态性状测试方法，观察记录大菊39个主要质量性状的表现，计算各性状的表现式样在品种群内的分布比例，结果显示舌状花的重瓣性、花瓣曲反状况、叶片基部性状、托叶大小等32个性状的表现式样在品种群内分布极不均匀，接近或超过70%的品种集中在性状的1种或2种表现式样中；而花瓣形状、外花瓣表面色、外花瓣背面色、花中心部颜色、叶裂刻基部有无突起、茎的曲直性和茎色等7个性状的表现式样在品种群内的分布则相对均匀。4。

The results above showed:(1) The cultivars of large-flowered chrysanthemum are widely distributed in Beijing area with plentiful floral types and floral colors;(2) The cultivars with the rare types and colors are urgent for protection;(3) Most of the quantitative characters obeys to the normal distribution,some important traits agreed with skewed distribution.The probability grading can assist quantitative character in establishing new grading standards;(4) Most of qualitative characters is asymmetrical in cultivars,which might be caused by unbalanced differentiation of characters or linkages coming from emphasis characters selection;(5) The floral characters are relatively stable and have comparably large influence on classification,the characters of stipule also have a certain degree of stability and can act as an reasonable supplement for cultivars classification,but the foliar characters have worse stability and can only be recognized as lower level criterion.(6) The genetic relationship among the Flat-petaled group,the Quilled-petal group and the Spoon-petaled group are closest,which followed by the Filiform-petal group because of long-term selection pressure for the specialization of the petal tip loved by most people.The Anemone-petal group has further genetic relationship with these four groups.(7) ISSR molecular marker detection indicates that the proportion of polymorphic loci of lardge-flowered chrysanthemum is higher,in which the Flat-petaled group has the highest genetic diversity level.

综合以上的研究结果表明：(1)北京地区现有大量大菊品种，且花型与花色较为丰富；(2)珍稀花型、花色品种亟需保护；(3)大部分数量性状是符合正态分布的，少数重点观赏性状迫于选择压力呈现偏态分布，概率分级法能够辅助大菊的数量性状建立新的分级标准；(4)大部分质量性状在品种群内的分布是不均匀的，可能是由菊花性状遗传分化的不均衡性或受到选育重点性状的连锁作用而造成的；(5)大菊花部性状相对稳定，对品种分类影响较大，其托叶部分也具有一定程度的稳定性，可以作为品种分类的合理补充，而叶部性状稳定性较差，在品种分类中应作为更次一级的分类依据；(6)平瓣类、管瓣类和匙瓣类的亲缘关系最近，畸瓣类品种花瓣先端特化受到人们的喜爱，长期的选择压力使其区别于3种基本瓣型，而这四种瓣型与桂瓣类在亲缘关系上较远；(7)ISSR分子标记检测表明：大菊品种多态性位点百分率较高，平瓣类品种的遗传分化最为丰富。

Leaves opposite or rarely alternate, subequal in size; stipules lanceolate, 7-10 mm; petiole 1.5-6(-10) cm, densely pubescent; leaf blade dark green or black when dry, oblate to 5-angled or oblate to orbicular-ovate, often ovate in upper stems, 8-12(-18)× 7-14(-22) cm, papery, secondary veins 2 pairs along midvein, abaxial surface densely pubescent along veins and veinlets, adaxial surface roughish, densely strigose, base truncate, subcordate, or broadly cuneate, margin coarsely 8-12-dentate, teeth gradually larger distally, often biserrate apically, apex 3-cusped or lobed, tip serrulate. Glomerules on axillary spikelike flowering branches, male branches often branched, 8-17 cm; female spikes distal, unbranched or branched, 5.5-24 cm.

叶对生或很少互生，大小近等；托叶披针形，7-10毫米；叶柄1.5-6(-10)厘米，密被短柔毛；叶片深绿色或黑色的干燥时，扁球形到具5个角或扁球形的到圆形卵形的，通常在上面茎内卵形的， 8-12(-18)* 7-14(-22)厘米，纸质，次脉2 对沿中脉，背面密被短柔毛的沿脉和细脉，正面微糙，密被糙伏毛，基部截形，近心形，或宽楔形，团伞花序在腋生的穗状的花枝，雄性分枝通常分枝，8-17厘米的；雌性穗状花序上部，不分枝或分枝，-24 厘米。

Stipules (2-)3-5 mm; petiole 3-5 mm, sulcate adaxially; leaf blade narrowly lanceolate or narrowly elliptic, 7-15 × 1.5-3 cm, chartaceous, glabrous on both surfaces, base narrowly cuneate, margin with dense and unequal glandular teeth, apex acuminate; midvein prominent on both surfaces; secondary veins numerous,± parallel; veinlets

托叶(2-)3-5毫米；叶柄3-5毫米，正面具槽；叶片狭披针形的或狭椭圆形， 7-15 * 1.5-3 厘米，纸质的，两面无毛，基部狭楔形，边缘具紧密的和不等长的具腺的齿，先端渐尖；中脉的两面突出；次脉多数，多少平行；细脉

Stipules green, falcate, rarely ovate or ovate-lanceolate, herbaceous, margin sharply serrate or lobed, rarely entire, apex acute or acuminate; petiole sparsely pilose or pubescent; leaf blade interrupted imparipinnate with (2 or)3 or 4 pairs of leaflets, reduced to 3 leaflets on upper leaves; leaflets sessile or shortly petiolulate, obovate, obovate-elliptic, or obovate-lanceolate, 1.5–5 × 1–2.5 cm, abaxially appressed pilose on veins, or densely pubescent or tomentose-pubescent between veins, rarely glabrescent, markedly or sparsely glandular punctate, adaxially pilose, or hirsute or hirtellous on veins, rarely glabrescent, base cuneate to broadly so, margin acutely to obtusely serrate, apex rounded to acute, rarely acuminate. Inflorescence terminal, spicate-racemose, branched or not; rachis pilose.

绿色，镰刀形的托叶，卵形或卵状披针形的很少，草本，边缘锐切成锯齿状或者有叶，很少全缘，先端锐尖或渐尖；叶柄疏生柔毛或短柔毛；叶片间断奇数羽状具(2或)3或4小叶，退化至3小叶在上部叶片上；短的小叶无柄或具小叶柄，倒卵形，倒卵状椭圆形，披针形或倒卵形， 1.5-5 * 1-2.5 厘米，背面的贴伏的具柔毛的在脉上，在脉之间的或密被短柔毛的或被绒毛的短柔毛，很少脱落无毛显著具点，正面具柔毛的或疏生腺体，或具粗毛或在脉上，很少脱落无毛，基部楔形到宽楔形，花序顶生，穗状总状，与否的分枝；轴具柔毛。

Such plants usually have one or more domatia, cavities in stems, leaves or roods that house ants. These may be elaborate systems of cavities with many chambers, e.g. Myrmecodia, or simple hollows in internodes, leaf veins, petioles or stipular spines, e.g. whistling thorns.

这类植物通常都具有一个或多个虫菌穴，虫菌穴是在植株的根、茎、叶上用来容纳蚂蚁的空腔，这些空腔具有许多发达的小室，例如茜草科具有简单而中空的结间部，阿拉比卡咖啡具有含有空腔的叶脉，有的植物叶柄及托叶处也有这样的结构，例如牛角刺槐。

Leaves alternate; stipules lanceolate, 4-8 mm; petiole 2-11(-13) cm, pubescent or glabrous; leaf blade ovate, elliptic, oblong or subelliptic,(5-)9-21 × 2.5-10 cm, papery, slightly rough, secondary veins 3 or 4 pairs along midvein, abaxial surface strigillose or pubescent on veins, rarely glabrous, adaxial surface sparsely appressed strigillose or glabrous, base broadly cuneate, rounded, or obtuse, margin denticulate, apex acuminate or caudate-acuminate. Glomerules usually unisexual, sometimes bisexual, in the axils of current and fallen leaves along normal leafy stems, 4-10 mm in diam.; male ones on proximal part of flower-bearing branches; female ones on distal part.

叶互生 托叶披针形，4-8毫米；叶柄2-11(-13)厘米，被短柔毛或无毛；叶片卵形，椭圆形，长方形或，( 5-)9-21 * 2.5-10 厘米，纸质，粗糙的稍，次脉3或4 对沿中脉，背面具短糙伏毛的或短柔毛在脉上，很少无毛，正面疏生贴伏具短糙伏毛的或无毛，通常单性的团伞花序，有时两性，在正常生叶的茎的叶和脱落叶的腋内，直径4-10毫米；在忍受花的分枝的下部上的雄性的；在上部上的雌性的。

Stipules gradually deciduous, triangular, 1-2 mm, membranous; petiole often unequal in length, 0.5-7 cm; leaf blade dark green or green blue, ovate, lanceolate, or obovate-oblong, asymmetric, unequal or equal in size, 1-15 × 0.6-5 cm, thinly papery or submembranous, 3(or 5)-veined, lateral veins many, irregularly reticulate, external secondary veins anastomosing by margin, sometimes inconspicuous, cystoliths finely fusiform, conspicuous adaxially, base obliquely rounded or subcordate, sometimes cuneate, margin entire or sometimes undulate, apex shortly or long caudate-acuminate. Inflorescences solitary; male inflorescence a cymose panicle or racemelike, often longer than leaves; female cymes as long as or shorter than leaves, pedunculate.

逐渐落叶，三角形的托叶，1-2毫米，膜；叶柄通常不等长，0.5-7厘米；蓝色的深绿色的叶片或绿色，卵形，，或倒卵形长圆形，不对称，不等长或大小相等， 1-15 * 0.6-5 厘米，薄纸质的或近膜质， 3(或5)脉，侧脉很多，不规则网状，外部次脉网结的在边缘，有时不明显花序单生；雄花序一总状花序状的密伞圆锥花序或，通常长于叶；雌性聚伞花序等长或短于叶，有花序梗。

Leaves simple; stipules subulate; leaf blade ovate-triangular or palmately 3-5-lobed or -parted. Flowers axillary, solitary or fascicled, sometimes grouped into terminal racemes.

单叶 托叶钻形叶片卵状三角形的或掌状的3-5浅裂或深裂花腋生的，单生或簇生，有时组合成顶生的总状花序的。

But we don't care about Battlegrounds.

但我们并不在乎沙场中的显露。

Ah! don't mention it, the butcher's shop is a horror.

啊！不用提了。提到肉，真是糟透了。