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Secondly, according to the stem and leaf anatomy, leaf epidermis, pollen morphology and floral development, it is found that there are no close relations between Acorus and Typhales, especially in the aspect of floral development. In the course of floral development of A. calamus, neither bracteloe primordium nor common primordium occurs in the whole course. Early development of the flower is strongly unidirectional. Three shield carpels develop into a pistil. In Sparganium fallax, each female flower has a bracteole. The bracteole primordium and the floral primordium originate from a complex structure. A single saccate carpel develops into a pistil. In staminate flowers, petal and stamen primordia initiate spirally.

菖蒲属与香蒲目的茎叶解剖特征、叶表皮特征、花粉特征及花器官发生特点等亦存在着明显区别,特别是花器官发生特点差异尤为显著,如菖蒲属花器官发生过程中,没有苞片原基,也不存在任何形式的复合原基,花被与雄蕊原基均以单向性方式发生,雌蕊由3心皮发育而成,具有"盾状心皮"的特点等,黑三棱属则有苞片原基发生,且苞片原基与雌花原基起源于共同的复合结构,雄花的花被原基和雄蕊原基均以螺旋状方式发生,雌蕊由单心皮发育而成,未发现有2或3个心皮原基或其痕迹的现象,体现出"囊状心皮"的特点。

Secondly, according to the stem and leaf anatomy, leaf epidermis, pollen morphology and floral development, it is found that there are no close relations between Acorus and Typhales, especially in the aspect of floral development. In the course of floral development of A. calamus, neither bracteloe primordium nor common primordium occurs in the whole course. Early development of the flower is strongly unidirectional. Three shield carpels develop into a pistil. In Sparganium fallax, each female flower has a bracteole. The bracteole primordium and the floral primordium originate from a complex structure. A single saccate carpel develops into a pistil. In staminate flowers, petal and stamen primordia initiate spirally.

菖蒲属与香蒲目的茎叶解剖特征、叶表皮特征、花粉特征及花器官发生特点等亦存在着明显区别,特别是花器官发生特点差异尤为显著,如菖蒲属花器官发生过程中,没有苞片原基,也不存在任何形式的复合原基,花被与雄蕊原基均以单向性方式发生,雌蕊由3心皮发育而成,具有&盾状心皮&的特点等,黑三棱属则有苞片原基发生,且苞片原基与雌花原基起源于共同的复合结构,雄花的花被原基和雄蕊原基均以螺旋状方式发生,雌蕊由单心皮发育而成,未发现有2或3个心皮原基或其痕迹的现象,体现出&囊状心皮&的特点。

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