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This paper has studied the physiological and biochemical changes during the period of forming strong buds of tree peony, and expect to provide academic basis for extending its view value and economic benefit . The main results are as the following:1 Under the field condition, Characteristics of photosynthesis of tree peony are very regular. From before blooming to the prophase of abducting buds that after blooming ,that is April,May and June,the net photosynthetic rate of tree peony is higher than the other period .It indicates that this period is very important for accumulating photosynthetic products of tree peony . So, if we want to get quality flower of tree peony ,it is very important to strengthen the management of tree peony cultivation and to ensure provide the fertilizer and water.2 From the middle ten days of June, with the time elapsing and with the environment conditions worsening, the characteristics of photosynthesis of tree peony decline deeply. But it is different among different tree peony breeds. For example , cv. Wu Long Peng Sheng can also has higher net photosynthetic rate, it indicates that cv. Wu Long Peng Sheng is super than the other two tree peony breeds when adapting the environment conditions and cv. Wu Long Peng Sheng has wide prospect to extend .3 The main factors of effecting the characteristics of photosynthesis are stomatal factor and non-stomatal factor ,that is the ability of photosynthesis of leaf cells. Under the environment of high light intensity in midsummer, if overshadow tree peony properly, then it can slow the declining of chlorophyll content, and increase the net photosynthetic rate and benefit for the accumulating of photosynthetic products.4 The content of soluble sugar in the root of tree peony is the highest among different organs during the period of forming strong buds . According to analyse ,we believe that the soluble sugar will compose starch then, and provide the solid base of material for the growth and development next year. It provide us strong academic basis of adopting the method of cultivating root and protecting root under thefield environment conditions .In other words ,it will help to increase the health level of the whole plant of tree peony and will be beneficial to forming quality tree peony flower if we strengthen the management of root. 5 To tree peony, the endogenesis hormone is important for its growth and development .In the prophase ,that is about the blooming of tree peony and the mid-summer ,IAA and ZR contents are higher and GA content is lower and its changes are very little, and the content of ABA is decreasing . This indicates that IAA and ZR act the important promote function for growth and development of tree peony in this period, and the function of GA is not important. Bur in the period of anniversary senescence of tree peony , the content of IAA and ZR begin to decrease ,while the ABA content has a certain rebound .T

本研究对几个牡丹品种壮芽形成过程中的某些生理生化变化做了研究,以期了解牡丹生长发育过程中生理生化的变化规律,探讨大田栽培条件下提高牡丹花质量的栽培措施和管理技术,为提高牡丹的观赏价值和经济效益提供理论依据,结果如下:1 大田栽培条件下,牡丹光合特性在生长发育过程中表现出极强的规律性,牡丹各品种的光合速率从牡丹开花前,到牡丹花谢后的花芽诱导期前期(6月10日之前),也就是在4、5、6月份,各牡丹品种的净光合速率相对与其他时间都维持在比较高的水平,这说明,在这段时间,是牡丹光合产物积累的重要时期,因此,要保证得到高质量的牡丹花,在这段时间加强对牡丹的栽培管理,保证这个时期的肥水供应十分重要。2 从6月中旬后,随时间的推移,环境条件的恶化,牡丹的光合性能剧烈下降,但是不同牡丹品种间仍有差异,比如,乌龙捧盛仍可维持较高的光合水平,这说明了乌龙捧盛这个牡丹品种对于环境的适应性较之其他牡丹品种高,有着广泛的推广前景。3 影响牡丹光合性能的主要因子是气孔因素和非气孔因素即叶肉细胞的光合能力,在盛夏高温高光强的环境条件下,对牡丹进行适当遮荫,能缓解牡丹叶绿素水平的下降,适当提高Pn,有利于光合产物的积累。4 可溶性糖在牡丹各品种的壮芽形成过程中,根中可溶性糖的含量在牡丹各部位中始终维持最高水平,分析认为这些可溶性糖在根系中进一步合成淀粉等储藏性物质,为来年牡丹的生长提供雄厚的物质基础,这为我们在大田栽培条件下对牡丹采取养根护根的栽培措施提供了有力的理论依据,即在大田栽培条件下,加强对牡丹根系的管理,有助于提高牡丹整株的健壮程度,有益于形成高质量的牡丹花。5内源激素对牡丹的生长发育调节作用是十分明显的,在前期,即开花前后和盛夏,IAA和ZR含量维持在比较高的水平,GA含量低且变化不明显,ABA含量逐渐降低,说明了IAA和ZR对牡丹在这段时间内的生长发育起到重要的促进作用,GA的作用不是很明显,ABA可能在高温高光强下有明显的升高,也说明了ABA的确有一些促进作用,但是到了牡丹周年衰老期,IAA和ZR的含量开始下降,ABA含量却有了一定程度的反弹,说明了IAA和ZR的作用开始降低,ABA起到了促进衰老的作用

There remained the generic conditions imposed by natural, as distinct from human law, as integral parts of the human whole: the necessity of destruction to procure alimentary sustenance: the painful character of the ultimate functions of separate existence, the agonies of birth and death: the monotonous menstruation of simian and human females extending from the age of puberty to the menopause: inevitable accidents at sea, in mines and factories: certain very painful maladies and their resultant surgical operations, innate lunacy and congenital criminality, decimating epidemics: catastrophic cataclysms which make terror the basis of human mentality: seismic upheavals the epicentres of which are located in densely populated regions: the fact of vital growth, through convulsions of metamorphosis, from infancy through maturity to decay.

324截然不同于人为的法则,这里依然存在着按照自然的法则作为对维持整个人类的生存不可分割的部分加诸于人的生物学之基本条件。为了获得有营养的食品,就不得不进行破坏性的杀戮。孤立的个人生存中终极机能那充满了苦恼的性质。生与死的痛苦。类人猿和人类女性那单调的月经,自初潮期一直延续到闭经期。海洋上、矿山和工厂里那些不可避免的事故;某些非常痛苦的疾病以及伴随而来的外科手术;生来的疯颠,先天性犯罪癖;导致人口大批死亡的传染病;在人类心灵深处种下恐怖种子的灾难性特大洪水;震中位于人口密集地区的大地震;历经剧烈变形,自幼年经过成熟期进入衰退期的生命成长的事实。

Comparative study on two levels of population and modules of mutualspecies in the two stations demonstrates that feature index tend to diminish suchas height,tussock amplitude,size,number,density and dry weight;and thegrowth analysis index incline to increase such as RGR、ULA、chlorophyllcontent and effectiveness of plant multi-leaves of the populations and DULA、DUMR、DAGA and DRGRa of the modules;and the dynamic curves of thepopulations and the modules also tend to steadily increase along with thetemperature ascending,heat accumulation and combination between water andheat in moderate-temperature steppe from fierce fluctuation alongside naturalprecipitation in warm-temperature steppe,illustrating that the transformabletendency from increasing the numbers of tussocks and tillers in order to expandthe resource range to seize in warm-temperature steppe to steadily improve drymatter production function of the populations and the modules in moderate-temperature steppe.

两个不同热量型研究站点共有种种群与构件两个水平的比较研究表明:随纬度增加和气温降低,种群与构件高度、丛幅、大小、数量、密度和干重等特征指标趋于降低和减少,而种群RGR、ULA、叶绿素含量和植物多叶性与构件DULA、DUMR、DRGR和DRGRa等生长指标则趋于增加,种群与构件的生长动态曲线也从随天然降水而剧烈波动趋向于随气温升高、热量积累和水热配置状况而稳定增长,说明各共有种从暖温型草原区增加植物种群与构件的大小和数量以扩充资源利用范围向中温型草原稳定地提高种群与构件的干物质生产性能的方面转变的趋势;同时,物质与能量的分配也从较多地投资于繁殖器官产生大量后代以适应未来降水的不可预期变化趋向于同化系统的构建和干物质生产效能的提高以抵御低温和干旱对物种生存的威胁,充分显示出不同温度条件及其水热组合对植物种群形态结构和生长繁育模式的显著影响及其响应特征。

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