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During the postgenomics era the information about genomic sequence and gene functions provides a new foundation for evolutionary biology and ecology As the first whole-genome sequenced plant Arabidopsis thaliana and its wild relatives have played a critical role in understanding the evolution of genomics and speciation Both A halleri and A lyrata are closely related to the model species A thaliana A halleri ssp gemmifera occurs in northeastern China Japan and Taiwan; while its sister A halleri ssp halleri is mainly distributed in Europe Geographical barriers such as Tienshan Mountain Range isolate these intraspecific sisters Likewise A lyrata ssp kamchatica and ssp lyrata occur in East Asia and North America respectively Such distribution patterns seem to be consistent with allopartic speciation The comparison between ancestral and extant polymorphism by multilocus can be informative about the population genetics of speciation In this study we collected and analyzed DNA sequences of 98 genes from four wild relatives of A thaliana A halleri ssp gemmifera A halleri ssp halleri A lyrata ssp kamchatica and A lyrata ssp lyrata The ancestral states of these four species were compared to each other in terms of level of genetic variation However the ancestral species at the time of speciation were substantially more polymorphic than the extant geographical populations The observations are not fully compatible with speciation by strict allopatry At some species pairs parapatric speciation seems more reasonable in speciation of Arabidopsis The 98 gene sequences are also used for the congruence test between gene genealogy and species phylogeny Only 28 genes support the species phylogeny but there are 23 genes supports another major genealogy { lyrata} thaliana Based on the phylogenetic position change of A lyrata ssp kamchatica and Ks value for each species pair suggested the recent directional gene flow between A halleri ssp gemmifera and A lyrata ssp kamchatica
阿拉伯芥是第一个完成基因体定序的开花植物,其基因体资讯提供植物学研究的重要依据;在解析阿拉伯芥属物种的亲缘关系以及种化机制等重要的演化议题时,阿拉伯芥近缘的野生物种自然成了不可或缺的关键;跟阿拉伯芥近缘的物种包括A halleri及A lyrata,其中A halleri ssp gemmifera主要分布於中国东北、日本以及台湾,与近缘的A halleri ssp halleri其分布於欧洲隔著天山及大陆的障蔽,而A lyrata ssp kamchatica主要分布於东北亚及台湾,与分布於北美五大湖的A lyrata ssp lyrata被北极圈所分隔,这样的分布模式暗示异域种化的可能。藉由多基因分析比较祖先物种与现生物种遗传歧异度的相关可提供讯息探讨种化时期的族群遗传结构,本研究针对A halleri ssp gemmifera、A halleri ssp halleri及A lyrata ssp kamchatica、A lyrata ssp lyrata四个物种,两对互为亚种的姊妹群,以阿拉伯芥为外群进行研究,在四个物种完成98个同源基因的分子序列,利用套装软体MCMCcoal来估算祖先物种的遗传变异,亦估算现生物种的核苷酸歧异度,观察到?多物种配对中祖先物种遗传多型性大於现生物种DNA歧异度,显示异域种化模型并无法完全解释阿拉伯芥属物种的种化模式,在某些物种配对间邻域种化模式应比异域种化更为可能;在基因树与物种树的比较,98个基因片段的亲缘模式只有28个是与已知物种树一致的,有23个基因其树状图支持{ lyrata} thaliana的型式,藉由kamchatica位置的变化以及估算各物种配对间的平均同义置换率,推测在A halleri ssp gemmifera与A lyrata ssp kamchatica间具有近代的单方向基因交流。
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Sampling error or allowable error is conventionally accepted between 1% and 10%; however, it is suggested that sampling error should be carefully assessed in light of the nature of the derivation of the population proportion and other aspects of the initial data.
抽样误差或允许误差是接受常规的1 %和10 %,但是,有人建议,抽样误差应当认真评估根据的性质推导的人口比例和其他方面的初步数据。
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The object of projectile design is analyzed, and the layer-TOPSIS method is provided for the evaluation of total effectiveness by taking the layer concept into the TOPSIS method. In the calculation of object weights, the grade calculation method is suggested, which has less dependence on the subjectivity of decision-maker and can take account of both sensitivity and allowableness of objects, the geometric-arithmetic method is provided for grade weight calculation and methods for sensitive weight and allowable weight are given. Of modes in system analyzing, the penetration model of target with spherical-fragment is carried out based on the theory of Dynamic Spherical Cavity Expansion, and it is a supplement of the models for calculation the damage of the fragmentation warhead to target.
对弹丸设计进行了目标分析;通过在TOPSIS方法中引入层次结构的概念,提出了适用于弹丸总体效能评价的层次TOPSIS评价方法;在指标权重计算中,提出了对决策者主观依赖较少的能反映指标的敏感性和相容性的分级权重计算方法;在分级权重计算中,分别给出了等比-等差权值计算方法和敏感-相容权值计算方法;在系统分析的模型中,对破片战斗部的威力计算中的模型进行了补充,在空穴膨胀理论的基础上提出了球形破片侵彻有限厚靶板的侵彻计算模型。
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Whats more, they suggested widening the scope of the almsman and building the new charities.
此外,他们还主张慈善组织扩大救助对象,创建新的慈善机构。
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They didn't allow us to make the alterations suggested.
他们不允许我们作提出的修改。
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Incorrect: The black and blue mark suggested that he had been involved in an altercation.
复合形容词由两个或两个以上的形容词组成用以修饰一个名词。
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It suggested that the over production of NO promoted the secretion of TNFα by alveolar macrophages.
这表明由于大量NO的产生促进了TNFα的分泌。
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Since microspore mother cells showabortion during the first meiotic divison,it is suggested thatthe early male abortion is frequently associated with variousdegrees of impairment to the fertility to the female〓amate.
研究的27个杂种第一代中,有11个组合的产量超过对照或与对照持平,占总组合数的40.74%。在强优势组合中,不育细胞质带来的负效应不足以使F1产量的显著降低,而且通过杂优补偿作用可比对照有所提高。
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It is suggested that the stratum intermedium participate in the differentiation of ameloblast and formation of enamel.
中间层细胞的增殖和凋亡及其TNSAP的表达与成釉细胞具有相关性,提示中间层细胞有可能参与成釉细胞的分化及釉质形成。
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On the basis of analyzing limitation of using penalty function method dealing with constraints , an amending method based on the knowlege regulating strategy is suggested to amende the mapping relationship of infeasible constraints in decoding , thus making the regulated individuals map into the space to obtain the most promising optimal solution .
在分析惩罚函数法对约束处理效率较低的情况下,提出了一种基于知识调整策略的修正法,对解码中不满足约束的映射关系进行修正,使调整后个体映射到最有希望获得最优解的空间中。
- 推荐网络例句
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The split between the two groups can hardly be papered over.
这两个团体间的分歧难以掩饰。
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This approach not only encourages a greater number of responses, but minimizes the likelihood of stale groupthink.
这种做法不仅鼓励了更多的反应,而且减少跟风的可能性。
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The new PS20 solar power tower collected sunlight through mirrors known as "heliostats" to produce steam that is converted into electricity by a turbine in Sanlucar la Mayor, Spain, Wednesday.
聚光:照片上是建在西班牙桑路卡拉马尤城的一座新型PS20塔式太阳能电站。被称为&日光反射装置&的镜子将太阳光反射到主塔,然后用聚集的热量产生蒸汽进而通过涡轮机转化为电力