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B Of the six basic media MS, MS1/2 (half-strength of MS salts and vitamins), WPM, DKW, B5 and SH, MS1/2 was the most proper one to induce somatic embryos. Somatic embryos generally regenerated directly from excised zygotic cotyledons. PGRs combination affected somatic embryogenesis significantly. Medium with NAA 1mg/L, TDZ 0.05mg/L, IBA 2—10mg/L combined with BA 10mg/L, or IBA 10mg/L integrated with BA 0-2mg/L gave the highest induction rate. Excised zygotic hypocotyls had the strongest potential to produce callus. Callus induction was also affected significantly by media and PGRs. The proper callus induction condition was MS1/2 medium containing NAA 1mg/L, IBA 10mg/L, BA 2-5mg/L and TDZ 0.05mg/L. Harvest period affect somatic embryogenesis significantly. Zygotic embryo explants collected from the end of July to the middle of August had strong potential to generate somatic embryos, when endosperm finished solidification, different parts of the embryos were completely formed, the size of embryos occupied about 2/3 of the embryo sac. Provided with optimized conditions, direct somatic embryogenesis rate can attain to 33. 68%, and callus induction rate of hypocotyls was up to 90.7%. Cytological observation on megasporogenesis and zygotic embryogenesis of Manchurian ash showed that the ovary was twicarpellum, twilocular with two ovules each loculus. The ovule was tenuinucellar and anatropous, with one megasporcocyte. The development of embryo sac is of the Polygoum type.

体细胞胚胎发生研究的结果表明：(1)成熟过程中的合子胚是诱导水曲柳体细胞胚胎发生的最佳外植体材料；(2)在所试验到的MS、MS1/2（将MS的所有成分均减半）、WPM、DKW、B〓、SH等六种基本培养基中，MS1/2是最适合诱导水曲柳体细胞胚胎发生的基本培养基；(3)水曲柳的体细胞胚胎发生以直接发生为主，体细胞胚主要来自于从合子胚分离的完整子叶；(4)培养基中的激素组合对水曲柳的体细胞胚胎发生有显著影响，诱导直接体细胞胚发生较好的激素组合有NAA 1mg/L+IBA 2，5，10mg/L+BA 10mg/L+TDZ 0.05mg/L和NAA 1mg/L+IBA 10mg/L+BA 0，2mg/L+TDZ 0.05mg/L；(5)合子胚分离的下胚轴具有最强的愈伤组织诱导潜力，少数愈伤组织可以分化出体细胞胚；(6)愈伤组织的诱导也受培养基和激素配比的显著影响，最适宜诱导的培养条件为MS1/2+NAA 1mg/L+IBA 10mg/L+BA 2，5mg/L+TDZ0.05mg/L；(7)采种时间对体细胞胚胎发生有显著影响。7月末到8月中旬的合子胚具有较强的体细胞胚发生潜力，此时种子尚未成熟，胚乳已呈固态，种胚的各个部分已分化完全，种胚体积占胚腔的大约2/3；(8)在各自综合的最适条件下，完整子叶的体细胞胚诱导率可达33.68%，下胚轴的愈伤组织诱导率可达90.7%。

The significance of man is not in what he attains, but rather in what he longs to attain.

一个人的意义不在于他的成就，而在于他所企求成就的东西。

The significance of man is not in what he attains,but rather in what he longs to attain.

人的意义不在于他有何成就，而在于什么是他所渴望成就的。

The significance of man is not what he attains,but rather in what he longs to attain.

人的意义不在于他有何成就，而在于什么是他所渴望成就的！

He was a Machiavellian, who would employ all means to attain to power.

他是一个马基雅维里式的人，会用一切手段来获取权力。

There is moment when art attain almost to the dignity of manual labour .

在某些时刻，艺术几乎要达到手工劳作的庄严地步。

It is possible for human being to attain correct cognition, management, maintenance and utilization of environment system and service function with reasonable marking under market economy, given that an appropriate relationship between input-outputs is provided.

经过全面价值标度的环境系统及其服务功能，在市场经济运行中，通过合理的投入-产出关系可以得到人类的正确认识、合理管理、维护与使用，从而奠定环境资源永续利用和人类可持续发展的基础。

It is an art and you can gather knowledge from various sources to attain mastery in this field.

它是一门艺术，你可以收集各种来源的知识，以达到掌握在这一领域。

When WTP attain to stable and mature level, the ecological consideration character coefficient is equal to 1 which is the ideal status.

从这个角度，森林生态效益的货币量的科学化和社会检验是当支付意愿水平达到一个稳定的、成熟水平时，即生态效益补偿的特征系数等于1的理想状态。

Only then is it possible to attain the maximum number of points

这是唯一拿高分的方法。

In the United States, chronic alcoholism and hepatitis C are the most common ones.

在美国，慢性酒精中毒，肝炎是最常见的。

If you have any questions, you can contact me anytime.

如果有任何问题，你可以随时联系我。