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Inflorescence spikelike, cylindrical, composed of spiny or bristly deciduous burrs arranged along an angular, often sinuous rachis; burrs sessile or with an obconical basal stipe, each composed of 1 or more sessile spikelets surrounded by an involucre of spines and bristles; bristles flexuous or more often spinous,± flattened, grooved on the outer face, united below, the degree of union varying from a small basal disk to a deep cupule, inner spines or bristles often ciliate around spikelets. Spikelets lanceolate, acute; glumes unequal, shorter than spikelet, lower sometimes suppressed; lower floret membranous, staminate or neuter; upper floret firmer, protogynous.
花序穗状，圆筒状，由落叶的毛刺沿着一具角安排的具刺的或具刚毛，通常波状的轴组成；毛刺无梗或具基部的倒锥状柄，每一个由1无梗或更多组成以总苞的刺和刚毛包围；或更多的刚毛曲折通常spinous，多少扁平，在外面上具凹槽，下面的合生，协会的度从一小的基部的花盘变化到一深的壳斗，小穗披针形，锐尖；不等长的颖片，短于小穗，降低有时压抑；更低的小花膜质，雄蕊花的或中性；坚定的上面小花，雌蕊先熟。
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Spikelets protogynous, strongly laterally compressed, floret 1, falling entire from pedicel; rachilla extension absent; glumes equal,± equaling and enclosing floret, broadly lanceolate to oblong, membranous to thinly leathery, prominently 3-veined, strongly keeled, keel usually ciliate, infrequently winged, lower margins almost free or connate for up to half their length, apex obtuse, acute or shortly awned; lemma broadly lanceolate to ovate, usually thinly membranous, keeled, obscurely 5-veined, smooth, glabrous, lower margins often connate, awned from lower back, apex truncate to acute; awn straight when short or geniculate when longer, column smooth, usually twisted at maturity, bristle scabrid; palea absent or very small.
雌蕊先熟的小穗，强烈侧面压扁，小花1，从花梗那里变为全缘；小穗轴扩展缺席；颖片等长，多少等长和包围小花，宽披针形的到长圆形，膜质到薄革质，显著3脉，强烈龙骨状，外稃宽披针形的到卵形，直的芒什么时候短或者膝曲长时间，顺利的柱子，通常成熟时扭曲，粗糙的硬毛；内稃缺席或者很小。
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Protogyny The maturation of the stigma before the anthers.
雌蕊先熟：指柱头先于花粉囊成熟的现象。
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proterandrous dichogamy：雄性器官先成熟
雄性器官先成熟 proterandrous dichogamy | 自末端开始的 proterminal | 雌蕊先熟花 proterogynous flower;protogynous flower
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proterogyny：雌蕊先熟的
proterogynous flower 雌蕊先熟花 | proterogyny 雌蕊先熟的 | proterozoic era 元古代
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proterogynous：雌蕊先熟的
proterogenesis 幼体发生 | proterogynous 雌蕊先熟的 | proterogynous flower 雌蕊先熟花
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proterogynous flower：雌蕊先熟花
proterogynous 雌蕊先熟的 | proterogynous flower 雌蕊先熟花 | proterogyny 雌蕊先熟的
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proterogynous flower;protogynous flower：雌蕊先熟花
自末端开始的 proterminal | 雌蕊先熟花 proterogynous flower;protogynous flower | 雌蕊先熟性 proterogyny
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protogynous：雌蕊先熟的
protocooperation 群团 | protogynous 雌蕊先熟的 | protogynous flower 雌蕊先熟花
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protogynous flower：雌蕊先熟花
protogynous 雌蕊先熟的 | protogynous flower 雌蕊先熟花 | protogyny 雌性先熟
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protohydrogen：原氢
protogynous 雌蕊先熟的 | protohydrogen 原氢 | protohymenium 原子实层